A small red-and-black butterfly poises statuesque above the purple blossom of this tall field-thistle. With its long sucker it probes industriously floret after floret of the crowded head, and extracts from each its wee drop of buried nectar. As it stands just at present, the dull outer sides of its four wings are alone displayed, so that it does not form a conspicuous mark for passing birds; but when it has drunk up the last drop of honey from the thistle flower, and flits joyously away to seek another purple mass of the same sort, it will open its red-spotted vans in the sunlight, and will then show itself off as one among the prettiest of our native insects. Each thistle-head consists of some two hundred separate little bell-shaped blossoms, crowded together for the sake of conspicuousness into a single group, just as the blossoms of the lilac or the syringa are crowded into larger though less dense clusters; and, as each separate floret has a nectary of its own, the bee or butterfly who lights upon the compound flower-group can busy himself for a minute or two in getting at the various drops of honey without the necessity for any further change of position than that of revolving upon his own axis. Hence these composite flowers are great favourites with all insects whose suckers are long enough to reach the bottom of their slender tubes.
The butterfly\'s view of life is doubtless on the whole a cheerful one. Yet his existence must be something so nearly mechanical that we probably overrate the amount of enjoyment which he derives from flitting about so airily among the flowers, and passing his days in the unbroken amusement of sucking liquid honey. Subjectively viewed, the butterfly is not a high order of insect; his nervous system does not show that provision for comparatively spontaneous thought and action which we find in the more intelligent orders, like the flies, bees, ants, and wasps. His nerves are all frittered away in little separate ganglia distributed among the various segments of his body, instead of being governed by a single great central organ, or brain, whose business it always is to correlate and co-ordinate complex external impressions. This shows that the butterfly\'s movements are almost all automatic, or simply dependent upon immediate external stimulants: he has not even that small capacity for deliberation and spontaneous initiative which belongs to his relation the bee. The freedom of the will is nothing to him, or extends at best to the amount claimed on behalf of Buridan\'s ass: he can just choose which of two equidistant flowers shall first have the benefit of his attention, and nothing else. Whatever view we take on the abstract metaphysical question, it is at least certain that the higher animals can do much more than this. Their brain is able to correlate a vast number of external impressions, and to bring them under the influence of endless ideas or experiences, so as finally to evolve conduct which differs very widely with different circumstances and different characters. Even though it be true, as determinists believe (and I reckon myself among them), that such conduct is the necessary result of a given character and given circumstances—or, if you will, of a particular set of nervous structures and a particular set of external stimuli—yet we all know that it is capable of varying so indefinitely, owing to the complexity of the structures, as to be practically incalculable. But it is not so with the butterfly. His whole life is cut out for him beforehand; his nervous connections are so simple, and correspond so directly with external stimuli, that we can almost predict with certainty what line of action he will pursue under any given circumstances. He is, as it were, but a piece of half-conscious mechanism, answering immediately to impulses from without, just as the thermometer answers to variations of temperature, and as the telegraphic indicator answers to each making and breaking of the electric current.
In early life the future butterfly emerges from the egg as a caterpillar. At once his many legs begin to move, and the caterpillar moves forward by their motion. But the mechanism which set them moving was the nervous system, with its ganglia working the separate legs of each segment. This movement is probably quite as automatic as the act of sucking in the new-born infant. The caterpillar walks, it knows not why, but simply because it has to walk. When it reaches a fit place for feeding, which differs according to the nature of the particular larva, it feeds automatically. Certain special external stimulants of sight, smell, or touch set up the appropriate actions in the mandibles, just as contact of the lips with an external body sets up sucking in the infant. All these movements depend upon what we call instinct—that is to say, organic habits registered in the nervous system of the race. They have arisen by natural selection alone, beca............