Viviparous quadrupeds, as a rule, present no great variety of form in the organ of smell. In those of them, however, whose jaws project forwards and taper to a narrow end, so as to form what is called a snout, the nostrils are placed in this projection, there being no other available plan; while, in the rest, there is a more definite demarcation between nostrils and jaws. But in no animal is this part so peculiar as in the elephant, where it attains an extraordinary and strength. For the elephant uses its nostril as a hand; this being the instrument with which it conveys food, fluid and solid alike, to its mouth. With it, too, it tears up trees, coiling it round their stems. In fact it applies it generally to the purposes of a hand. For the elephant has the double character of a land animal, and of one that lives in swamps. Seeing then that it has to get its food from the water, and yet must necessarily breathe, inasmuch as it is a land animal and has blood; seeing, also, that its excessive weight prevents it from passing rapidly from water to land, as some other sanguineous vivipara that breathe can do, it becomes necessary that it shall be suited alike for life in the water and for life on dry land, just then as divers are sometimes provided with instruments for respiration, through which they can draw air from above the water, and thus may remain for a long time under the sea, so also have elephants been furnished by nature with their lengthened nostril; and, whenever they have to traverse the water, they lift this up above the surface and breathe through it. For the elephant’s proboscis, as already said, is a nostril. Now it would have been impossible for this nostril to have the form of a proboscis, had it been hard and incapable of bending. For its very length would then have prevented the animal from supplying itself with food, being as great an impediment as the of certain oxen, that are said to be obliged to walk backwards while they are grazing. It is therefore soft and flexible, and, being such, is made, in addition to its own proper functions, to serve the office of the fore-feet; nature in this following her wonted plan of using one and the same part for several purposes. For in polydactylous quadrupeds the fore-feet are intended not merely to support the weight of the body, but to serve as hands. But in elephants, though they must be reckoned polydactylous, as their foot has neither cloven nor solid hoof, the fore-feet, owing to the great size and weight of the body, are reduced to the condition of mere supports; and indeed their slow motion and unfitness for bending make them useless for any other purpose. A nostril, then, is given to the elephant for respiration, as to every other animal that has a lung, and is lengthened out and endowed with its power of coiling because the animal has to remain for considerable periods of time in the water, and is unable to pass thence to dry ground with any rapidity. But as the feet are shorn of their full office, this same part is also, as already said, made by nature to supply their place, and give such help as otherwise would be rendered by them.

As to other sanguineous animals, the Birds, the Serpents, and the Oviparous quadrupeds, in all of them there are the nostril-holes, placed in front of the mouth; but in none are there any distinctly formed nostrils, nothing in fact which can be called nostrils except from a functional point of ............