Democritus of Abdera and certain others who have treated of respiration, while saying nothing definite about the lungless animals, nevertheless seem to speak as if all breathed. But Anaxagoras and Diogenes both maintain that all breathe, and state the manner in which fishes and oysters respire. Anaxagoras says that when fishes discharge water through their gills, air is formed in the mouth, for there can be no vacuum, and that it is by drawing in this that they respire. Diogenes’ statement is that, when they discharge water through their gills, they suck the air out of the water surrounding the mouth by means of the vacuum formed in the mouth, for he believes there is air in the water.

But these theories are untenable. Firstly, they state only what is the common element in both operations and so leave out the half of the matter. For what goes by the name of respiration consists, on the one hand, of inhalation, and, on the ot............