After this, as said already, the internal parts come into being before the external. The greater become visible before the less, even if some of them do not come into being before them. First the parts above the hypozoma are differentiated and are superior in size; the part below is both smaller and less differentiated. This happens in all animals in which exists the distinction of upper and lower, except in the insects; the growth of those that produce a scolex is towards the upper part, for this is smaller in the beginning. The cephalopoda are the only locomotive animals in which the distinction of upper and lower does not exist.

What has been said applies to plants also, that the upper portion is earlier in development than the lower, for the roots push out from the seed before the shoots.

The agency by which the parts of animals are differentiated is air, not however that of the mother nor yet of the embryo itself, as some of the physicists say. This is manifest in birds, fishes, and insects. For some of these are separated from the mother and produced from an egg, within which the differentiation takes place; other animals do not breathe at all, but are produced as a scolex or an egg; those which do breathe and whose parts are differentiated within the mother’s uterus yet do not breathe until the lung is perfected, and the lung and the preceding parts are differentiated before they breathe. Moreover, all polydactylous quadrupeds, as dog, lion, wolf, fox, jackal, produce their young blind, and the eyelids do not separate till after birth. Manifestly the same holds also in all the other parts; as the qualitative, so also the quantitative differentia comes into being, pre-existing potentially but being actualized later by the same causes by which the qualitative distinction is produced, and so the eyelids become two instead of one. Of course air must be present, because heat and moisture are present, the former acting and the latter being acted upon.

Some of the ancient nature-philosolphers made an attempt to state which part comes into being after which, but were not sufficiently acquainted with the facts. It is with the parts as with other things; one naturally exists prior to another. But the word ‘prior’ is used in more senses than one. For there is a difference between the end or final cause and that which exists for the sake of it; the latter is prior in order of development, the former is prior in reality. Again, that which exists for the sake of the end admits of division into two classes, (1) the origin of the movement, (2) that which is used by the end; I mean, for instance, (1) that which can generate, (2) that which serves as an instrument to what is generated, for the one of these, that which makes, must exist first, as the teacher before the learner, and the other later, as the pipes are later than he who learns to play upon them, for it is superfluous that men who do not know how to play should have pipes. Thus there are three things: first, the end, by which we mean that for the sake of which something else exists; secondly, the principle of movement and of generation, existing for the sake of the end (for that which can make and generate, considered simply as such, exists only in relation to what is made and generated); thirdly, the useful, that is to say what the end uses. Accordingly, there must first exist some part in which is the principle of movement (I say a part because this is from the first one part of the end and the most important part too); next after this the whole and the end; thirdly and lastly, the organic parts serving these for certain uses. Hence if there is anything of this sort which must exist in animals, containing the principle and end of all their nature, this must be the first to come into being — first, that is, considered as the moving power, but simultaneous with the whole embryo if considered as a part of the end. Therefore all the organic parts whose nature is to bring others into being must always themselves exist before them, for they are for the sake of something else, as the beginning for the sake of the end; all those parts which are for the sake of something else but are not of the nature of beginnings must come into being later. So it is not easy to distinguish which of the parts are prior, those which are for the sake of another or that for the sake of which are the former. For the parts which cause the movement, being prior to the end in order of development, come in to cause confusion, and it is not easy to distinguish these as compared with the organic parts. And yet it is in accordance with this method that we must inquire what comes into being after what; for the end is later than some parts and earlier than others. And for this reason that part which contains the first principle comes into being first, next to this the upper half of the body. This is why the parts about the head, and particularly the eyes, appear largest in the embryo at an early stage, while the parts below the umbilicus, as the legs, are small; for the lower parts are for the sake of the upper, and are neither parts of the end nor able to form it.

But they do not say well nor do they assign a necessary cause who say simply that ‘it always happens so’, and imagine that this is a first principle in these cases. Thus Democritus of Abdera says that ‘there is no beginning of the infinite; now the cause is a beginning, and the eternal is infinite; in consequence, to ask the cause of anything of this kind is to seek for a beginning of the infinite’. Yet according to this argument, which forbids us to seek the cause, there will be no proof of any eternal truth whatever; but we see that there is a proof of many such, whether by ‘eternal’ we mean what always happens or what exists eternally; it is an eternal truth that the angles of a triangle are always equal to two right angles, or that the diagonal of a square is incommensurable with the side, and nevertheless a cause and a proof can be given for these truths. While, then, it is well said that we must not take on us to seek a beginning (or first principle) of all things, yet this is not well said of all things whatever that always are or always happen, but only of those which really are first principles of the eternal things; for it is by another method, not by proof, that we acquire knowledge of the first principle. Now in that which is immovable and unchanging the first principle is simply the essence of the thing, but when we come to those things which come into being the principles are more than one, varying in kind and not all of the same kind; one of this number is the principle of movement, and therefore in all the sanguinea the heart is formed first, as was said at the beginning, and in the other animals that which is analogous to the heart.

From the heart the blood-vessels extend throughout the body as in the anatomical diagrams which are represented on the wall, for the parts lie round these because they are formed out of them. The homogeneous parts are formed by heat and cold, for some are put together and solidified by the one and some by the other. The difference between these has already been discussed elsewhere, and it has been stated what kinds of things are soluble by liquid and fire, and what are not soluble by liquid and cannot be melted by fire. The nutriment then oozes through the blood-vessels and the passages in each of the parts, like water in unbaked pottery, and thus is formed the flesh or its analogues, being solidified by cold, which is why it is also dissolved by fire. But all the particles given off which are too earthy, having but little moisture and heat, cool as the moisture evaporates along with the heat; so they become hard and earthy in character, as nails, horns, hoofs, and beaks, and therefore they are softened by fire but none of them is melted by it, while some of them, as egg-shells, are soluble in liquids. The sinews and bones are formed by the internal heat as the moisture dries, and hence the bones are insoluble by fire like pottery, for like it they have been as it were baked in an oven by the heat in the process of development. But it is not anything whatever that is made into flesh or bone by the heat, but only something naturally fitted for the purpose; nor is it made in any place or time whatever, but only in a place and time naturally so fitted. For neither will that which exists potentially be made except by that moving agent which possesses the actuality, nor will that which possesses the actuality make anything whatever; the carpenter would not make a box except out of wood, nor will a box be made out of the wood without the carpenter. The heat exists in the seminal secretion, and the movement and activity in it is sufficient in kind and in quantity to correspond to each of the parts. In so far as there is any deficiency or excess, the resulting product is in worse condition or physically defective, in like manner as in the case of external substances which are thickened by boiling that they may be more palatable or for any other purpose. But in the latter case it is we who apply the heat in due measure for the motion required; in the former it is the nature of the male parent that gives it, or with animals spontaneously generated it is the movement and heat imparted by the right season of the year that it is the cause.

Cooling, again, is mere deprivation of heat. Nature makes use of both; they have of necessity the power of bringing about different results, but in the development of the embryo we find that the one cools and the other heats for some definite purpose, and so each of the parts is formed; thus it is in one sense by necessity, in another for a final cause, that they make the flesh soft, the sinews solid and elastic, the bones solid and brittle. The skin, again, is formed by the drying of the flesh, like the scum upon boiled substances; it is so formed not only because it is on the outside, but also because what is glutinous, being unable to evaporate, remains on the surface. While in other animals the glutinous is dry, for which reason the covering of the invertebrates is testaceous or crustaceous, in the vertebrates it is rather of the nature of fat. In all of these which are not of too earthy a nature the fat is collected under the covering of the skin, a fact which points to the skin bein............